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- B.S., Biology, The University of Maryland at College Park, College Park, MD, (1988-1992)
- Ph.D., Pharmacology, The University of North Carolina, Chapel Hill, Chapel Hill, NC, (1992-1998)
- Postdoc, Developmental Biology, The Rockefeller University, New York, NY, (1998-2001)
- Postdoc, Developmental Biology, Memorial Sloan-Kettering Cancer Center, New York, NY, (2001-2006)